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Shell ?                                         SHELLS Links  pleuro.gifConvention
"It is perhaps a more fortunate destiny to have a taste for collecting shells than to be born a millionaire."

Shells are lovely natural objects, equals in beauty to any flower or butterfly, they are more than just pretty baubles found on beaches. They are the exterior skeletons (exoskeletons) of a group of animals called mollusks. The word "mollusk" means "soft-bodied;" an exterior skeleton is very important to these creatures, providing them with shape and rigidity, and also with protection, and sometimes camouflage, from predators.
Mollusks are classified into major groupings according to the characteristics of their shells.
Snails (Gastropoda) have a single shell which spirals outward and to one side as it grows. Most Cephalopoda (octopi and squid) have no shell, but the Chambered Nautilus of that group has a shell. This shell does coil, but it coils flatly, in a single plane. Tusk shells (Scaphopoda) also have a single shell, but it does not coil at all; it grows in a narrow and very slightly curved cone shape. Bivalves (Bivalvia), including oysters, clams, scallops and mussels, have two parts to their shells that enclose their tender bodies like the two halves of a hinged box. Chitons (Polyplacophora) are little armored tanks, with a row of eight overlapping plates protecting them. The Neopilina (Monoplacophora), are deep-sea "living fossils;" they have a single shell which hardly coils at all, but fits over their bodies like a protective cup. (Some gastropods (the limpets) have shells like this too, but their body structure is very different.) Last are the deepsea worm-like Aplacophora, with no shell at all, but little calcareous spines on their bodies. Mollusks make their shells from calcium they derive from their environment, either the food they eat or the water they dwell in. When a tiny mollusk hatches from its egg, it comes into the world equipped with a tiny shell. This shell is actually a part of the animal, growing as it grows, accommodating its needs. Each different species of mollusk makes a shell that is, in most cases, unique to it alone. Indeed, this uniqueness of form is partly what allows amateur shell collectors (conchologists) and professional scientists who study mollusks (malacologists) to determine a mollusk's species. Each species is destined genetically to develop the same type of shell its progenitors did. But, just as with humans, there are many distinct differences. Food, climate, environment, accident, and the mollusk's particular heredity all play their parts in making each shell an individual. Just as important as protection and rigidity, is the assistance a shell renders its maker in pursuit of the necessities of its life. Some shell shapes are streamlined for ease of burrowing through mud or sand. Some bivalves are heavily ridged in ways that help them stay anchored in the bottom where they live. Others are assisted in burrowing by special shell sculpture. There are shells that grow long spines to entrap and encourage growth of camouflaging seaweed and corals. Others, like cowry shells, are smooth and polished and highly glossy with bright interesting patterns on their shells. Often the animals themselves are patterned as well. When a predator disturbs the animal, it withdraws inside its shell, revealing a surface that is very different in color and pattern than the one the hungry predator had its eye on. It is speculated that quick change artistry has a startle effect on the predator. (And still other mollusks, like the slugs, squid and octopi, and those lovely creatures, the nudibranchs, have given up building a shell altogether, relying wholly on other defenses.) Indeed, a great deal may be learned about a mollusk's individual life by examining its shell. Many bear old healed-over breaks and chips that speak of a battle won with some predator. Others show color change which testifies to a change in diet or water chemicals. Mollusks that are very old in snail-years may become greatly thickened and dulled in color. Rough water conditions discourage the growth of long spines on normally spiny species, while quiet habitats allow extravagant spiny extensions. (It is now believed that this effect may have to do with a mollusk's different diet in rough water.) Sometimes a shell may have so great an encrustation of marine organisms on its shell that locomotion becomes impossible and the animal starves to death. And shells aren't found only on beaches. They can be found anywhere mollusks live. Both gastropods and bivalves inhabit rivers and streams and lakes, where they have made adaptations to freshwater living. Many species have also adapted to life on land, and can be observed in leaf litter, in trees, on plants, under rocks and buried in loose dirt. In the ocean, mudflats and mangrove areas are homes to hundreds of mollusk species. Some live on the tree roots and branches just at water line. Others bury themselves in the mud or sand bottom, emerging to feed, often at night or at low tide. There are mollusks which live only on coral reefs, and those which inhabit the ocean depths. Some, like the slit shell and the Neopilina, were thought to be extinct until they were rediscovered in these depths. Shells live on seaweed, on coral holdfasts, under rocks, buried in the roots of undersea grass beds. Some shells even float throughout their entire lives on a raft of bubbles. Each has a specialized habitat and an ecological niche.    

      Fabulous Shells


MARINE

When we think of shells, it is seashells that probably come to mind first, in a multiplicity of shapes, colors and patterns. Scallops, cowries, conchs and cones in rainbow hues. Lacy frills, elongate spikes, gleaming egg shapes, huge bowls and tiny rice grains, all characterize marine mollusks. Mollusks first evolved in the sea and have been adapting to its changing ecological niches for nearly 600 million years. So it is not surprising that marine species exhibit the phylum's greatest diversity. Mollusca in general exhibit more morphological diversity than any other phylum, including arthropods. The vast difference between the giant squid and microscopic nuculid clams is unmatched in the animal kingdom. All seven of the existing classes of the Mollusca are present in the ocean; only the Bivalvia (clams) and Gastropoda (snails) have moved into freshwater; Gastropoda is the sole class to adapt to life on dry land. Invertebrates, animals without backbones, are commonly presented with a special survival problem -- soft bodies, easily snapped up by a hungry predator. The shells which mollusks have evolved function to help them with this difficulty by presenting a tough exterior to predators in search of a meal. It bears mention here that many marine mollusks get along fine without a shell, having adapted in other ways which help them escape being eaten, at least long enough to reproduce their kind. Squid are fast swimmers and many species travel in schools. The octopus jets out a blob of "ink" to distract and fool a predator. Many sea slugs are able to consume stinging cells of other animals like coral. They transfer these natural deterrents to their own body cells where they function to instruct predators to avoid their kind. But in general, the phylum has flourished through epochs of geologic time wearing hard shells that afford excellent protection against all but the most determined foe. Mollusks have also adopted an amazing array of life styles and habitats. Some groups are carnivores, some are strict vegetarians, others are scavengers or parasites or commensals; the bivalves, for the most part are sedentary filter feeders, but some are predacious. Marine mollusks burrow, creep, tunnel, float or swim. They make their homes in mud, sand, silt, coral grit, rock, shell, tidal pool or grass. With such a multitude of lifestyles, it is not surprising that any number of evolutionary experiments and accidents have proved successful, explaining in part the fabulous diversity of pattern, size, and form among marine mollusks. Pattern often functions as camouflage. Wide flaring lips on conchs can help protect extended, grazing soft parts. Elongate anterior canals can likewise protect a mollusk's vulnerable siphonal "nose," stretched out to detect food or foe. Extended teeth on some muricids function as pry bars to pop open the protective plates of their food, barnacles. Low, flattened shells on limpets can withstand wave pressure. The high spires on many species aid in streamlining. Convoluted apertures function to keep out would-be predators. Thickened ribs add strength to shells of many species without adding commensurate weight and "construction costs." Specialized slits on abalones and others send waste products away from their water intake. In some cases mollusks from many different lineages have produced similar shells, probably because of a similarity of lifestyles, and similar constraints on how to build shells using only a logarithmic spiral, using only what their ancestors gave them. It seems to most of us that we'll never become familiar with all the types of mollusks in the sea. And then, just when we begin to feel comfortable, we discover the entire array of microscopic mollusks waiting for us to discover them, or the hundreds of millions of years worth of extinct shells, preserved as fossils, that occur in our rocks and wash out in streams. Or we find that we'd like to narrow our scope somewhat by specializing in a particular group of mollusks. There's always something new to hold our interest, some fresh discovery, or new scientific advances leading to changes in the taxonomy of mollusks. Some new family of bivalves catches our attention, or the intricacy of chiton plates (Class Polyplacophora) fascinates us, or we decide we'd like to specialize in the tusk shells, Class Scaphopoda, for a while. And we're off again, on a new pursuit of the magnificent diversity that is marine mollusks.

FRESHWATER

Freshwater mollusks include members of the Gastropoda and Bivalvia. To our knowledge, no cephalopods, chitons, tusk shells, or other groups ever invaded the freshwater environment. These invasions have happened over and over again, at different times, with different groups. The result is a hodge-podge of very unrelated animals all living in that creek in your back yard. Freshwater snails, for example, are a mix of neritoids, cerithioids, rissoids, and others. Some are the result of marine invasions, others represent a return to an aquatic habitat by terrestrial forms. Those evolved from marine forms may retain an operculum and breathe with a gill. Those evolved from terrestrial ancestors usually lack operculae and breathe with a 'lung' comprised of mantle tissue. In North America, the more diverse groups include the Pleuroceridae, Lymnaeidae, Physidae, and Vivipariidae. Sizes range from large 'apple' snails the size of baseballs, to microscopic hydrobiids and bithyniids. Most live by grazing algae, often tenaciously clinging to stones in swift currents. Freshwater bivalves also are a melting pot of independent invasions. Our Recent forms include species derived from veneroids, trigonioids, mytiloids, and even cardioids. They range from the tiny fingernail clams to the massive freshwater mussel Megalonaias nervosa, which reaches a length of 11 inches and weighs several pounds. Many have evolved specialized life cycles including brooding their young and producing larvae parasitic on fishes and amphibians. Some are free-living, burrowing in the substrate with their foot. Others may possess byssal threads for some or all of their life. And a few cement themselves to objects in the manner of marine ostreids. Freshwater species are perhaps the most imperiled group of all mollusks. Confined to narrow ranges the width of a river or creek, they are easily extirpated by ecological disasters - manmade or otherwise. Over 50 freshwater mussels are considered endangered in the United States. Perhaps another two dozen have become extinct in the last 200 years. Most of the dozens of pleurocerid snail species that inhabited the southern rivers of North America are extinct. Many freshwater mollusk species have been reduced to pitifully small populations, their continued existence at the mercy of the vagaries of nature and the ignorance of man.

What is a Mollusk?  

The phylum Mollusca includes organisms such as clams, snails, slugs, octopuses, squid, and chitons. As the phylum is currently defined, several features are common to all or most mollusks: A mantle that secretes calcium carbonate in the form of spicules or shell. A mantle cavity where respiration occurs, usually through the ctenidium (gill) in aquatic forms, or through the mantle wall in air in terrestrial ones, and where excretory and reproductive organs discharge. A body divided into three regions, the head, foot, and visceral mass Three coelomic spaces, for kidney, heart and gonad, A radula, a ribbon of teeth used in feeding. The word "mollusk" derives from Latin mollis meaning "soft," just as the term "malacology," the study of mollusks, comes from the Greek word for soft, malakos. Originally Mollusca was used for naked, soft-bodied animals, whereas shelled animals were placed in the Testacea. In the early 1800s, Baron Georges Cuvier realized that gastropods, bivalves, scaphopods, and cephalopods belonged in one group, but he also included barnacles and brachiopods in the Mollusca, which have since been removed. The modern term "shellfish" refers to shelled mollusks and to crustaceans, which are members of the phylum Arthropoda. The term "conchology" is also used for the study of mollusks, it is sometimes applied to the study of shells alone.
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